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If they are both running *at the same time*, like in separate user sessions if you have more than one user logged in at once, then yes. They both store your wireless network configuration in different ways (knm in native KDE formats, nm-applet in GConf for GNOME), and thus only one can be the authoritative source for user-scope network information for NM.
In the long run we'll change that so that the active user session gets to own the correct DBus service, no matter whether that session is KDE or GNOME, but that's not the case yet.
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